Oryx /
Volume
47 /
Issue 03 /
Mon Jul 01 00:00:00 BST 2013,
pp 369 - 380
Copyright © Fauna & Flora International 2013 The online version of this article is published within an Open Access environment subject to the conditions of the Creative Commons Attribution licence <http://creativecommons.org/licenses/by/3.0/>.
DOI:
http://dx.doi.org/10.1017/S003060531200172X
(About DOI),
Published online: Tue May 14 00:00:00 BST 2013